The
Vespidae are a large (nearly 5000 species), diverse,
cosmopolitan family of
wasps, including nearly all the known
eusocial wasps (such as
Polistes fuscatus,
Polistes annularis, and
Vespula germanica) and many wasps. Each social wasp
colony includes a
queen and a number of female
workers with varying degrees of sterility relative to the queen. In temperate social species, colonies usually only last one year, dying at the onset of winter. New queens and males (drones) are produced towards the end of the summer, and after mating, the queens hibernate over winter in cracks or other sheltered locations. The nests of most species are constructed out of mud, but polistines and vespines use plant fibers, chewed to form a sort of
paper (also true of some stenogastrines). Many species are pollen vectors contributing to the
pollination of several plants, being potential or even effective pollinators, while others are notable predators of pest insect species.