Opiliones (; formerly
Phalangida) is an order of
arachnids commonly known as
harvestmen. , over 6,500 species of harvestmen have been discovered worldwide, although the total number of
extant species may exceed 10,000. The order Opiliones includes five suborders:
Cyphophthalmi,
Eupnoi,
Dyspnoi,
Laniatores, and the recently named
Tetrophthalmi. With the exception of Dyspnoi (which is restricted to
North America and
Eurasia), representatives of each can be found on every continent except
Antarctica. Well-preserved fossils have been found in the 400-million-year-old
Rhynie cherts of Scotland, and 305-million-year-old rocks in France, which look surprisingly modern, indicating that their basic body plan appeared very early on, and, at least in some taxa, has changed little since that time. Their
phylogenetic position within Arachnida is disputed: their closest relatives may be the mites (
Acari) or the
Novogenuata (the
Scorpiones,
Pseudoscorpiones and
Solifugae). Although superficially similar to and often confused with
spiders (order
Araneae), Opiliones is a distinct order that is not closely related to spiders within Arachnida. They can be easily distinguished from even long-legged spiders by their fused body regions and single pair of eyes in the middle of their
cephalothorax (spiders have an 'abdomen' that is separated from the cephalothorax by a constriction, as well as three to four pairs of eyes, usually around the margins of their cephalothorax).