Archosauriformes (
Greek for 'ruling lizards', and
Latin for 'form') is a clade of
diapsid reptiles that developed from
archosauromorph ancestors some time in the Late
Permian (roughly 250
million years ago). It was defined by
Jacques Gauthier (1994) as the clade stemming from the last common ancestor of
Proterosuchidae and
Archosauria; Phil Senter (2005) defined it as the most exclusive clade containing
Proterosuchus and Archosauria. These
reptiles, which include members of the
family Proterosuchidae and more advanced forms, were originally superficially crocodile-like predatory semi-aquatic animals about 1.5 meters (5 ft) long, with a sprawling elbows-out stance and long snouts. Unlike the bulk of their
therapsid contemporaries, the proterosuchids survived the
catastrophe at the end of the Permian, perhaps because they were opportunistic scavengers, perhaps because they could retreat into water to find respite from an overheated climate. Any such scenarios are hypothetical; what is clearer is that these animals were highly successful in their new environment, and evolved quickly. Within a few million years at the opening of the
Triassic, the proterosuchids had given rise to the
Erythrosuchidae (the first
sauropsids to totally dominate their environment), who in turn were the ancestors of the small agile
Euparkeriidae, from which a number of successfully more advanced families – the archosaurs proper – evolved rapidly to fill empty ecological niches in the devastated global system. The archosaurs include
Crocodylia,
dinosaurs,
birds, and a few extinct
orders.