The
Temnospondyli (from
Greek τέμνειν (
temnein, "to cut") and σπόνδυλος (
spondylos, "vertebra")) are a diverse
order of small to giant
tetrapods—often considered
primitive amphibians—that flourished worldwide during the
Carboniferous,
Permian, and
Triassic periods. A few species continued into the
Cretaceous. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including fresh water, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the
larval stage,
metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales, claws, and armor-like bony plates, that distinguish them from modern amphibians.