The order
Salviniales (formerly known as the
Hydropteridales and including the former
Marsileales) is an order of
ferns in the class
Polypodiopsida. Salviniales are all aquatic and differ from all other ferns in being
heterosporous, meaning that they produce two different types of
spores (megaspores and microspores) that develop into two different types of
gametophytes (female and male gametophytes, respectively), and in that their gametophytes are endosporic, meaning that they never grow outside the spore wall and cannot become larger than the spores that produced them. In being heterosporus with endosporic gametophytes they are more similar to
seed plants than to other ferns.