The
Protura, or
proturans, and sometimes nicknamed
coneheads, are very small (<2 mm long), soil-dwelling animals, so inconspicuous they were not noticed until the 20th century. The Protura constitute an
order of
hexapods that were previously regarded as insects, and sometimes treated as a
class in their own right. Some evidence indicates the Protura are
basal to all other hexapods, although not all researchers consider them
Hexapoda, rendering the
monophyly of Hexapoda unsettled. Uniquely among hexapods, proturans show
anamorphic development, whereby
body segments are added during
moults. Szeptycki (2007) lists a total of 731 described species worldwide, in seven
families, nearly 300 of which are contained in a single
genus,
Eosentomon.