Metatheria is one of two mammalian
clades, the other being
Eutheria, with extant members that diverged in the
Early Cretaceous or perhaps the
Late Jurassic, and which includes all
mammals more closely related to
marsupials than to
placentals. First proposed by
Thomas Henry Huxley in 1880, it is a slightly more inclusive group than the
marsupials; it contains all of the living mammals with abdominal
pouches (most female marsupials) as well as their more primitive ancestors and relatives. Metatherians are one of three main classes of extant mammals:
monotremes (egg laying mammals like the platypus and the echidna), metatheria (or marsupials, which includes the three American orders (Didelphimorphia, Paucituberculata and Microbiotheria) and the four Australasian orders (Notoryctemorphia, Dasyuromorphia, Peramelemorphia and Diprotodontia)), and the eutherians (or placental mammals) consisting of twenty one orders, divided into four superorders. Metatherians belong to a subgroup of the northern tribosphenic mammal clade or Boreosphenida. They differ from all other mammals in certain morphologies like their dental formula, which includes about five upper and four lower incisors, a canine, three premolars, and four molars. Other morphologies include skeletal and anterior dentition, such as wrist and ankle apomorphies; all metatherians share derived pedal characters and calcaneal features.